Tube rectifiers convert AC to DC and is used in the power supply of an amplifier. The 12at7 tubes are signal tubes in your preamp.
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Because the 12AT7 is directly in the signal path it's obviously going to change the sound. But don't dismiss the rectifier as they can also change the flavor of the sound.
Here is a thread about subs for the 5Z4p. I have never used this tube but I have used a couple of the subs mentioned.