I’ve got the HIFIMAN HE400S and find them to be very neutral and open sounding. There’s a review of the LCD-X on headphones.com that compares them to to the HIFIMAN Ananda (a couple models up from mine) and finds them pretty comparable sonically, and at around 400g they’re almost 40% lighter than the Audeze. As a bonus they’re “only” $700 but can be had open box for $600. The Arya would be the next step up at about double the price. Hope this helps.
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The most comfortable headphones I own (I have about 10 or 12) are my Sennheiser / Drop HD6XX - https://drop.com/buy/massdrop-sennheiser-hd6xx. Changing out the earpads can make them even more comfortable. Not just comfortable, but lots of bang for the buck. Maybe not quite on the same level as the LCD-X in terms of detail and bass quality/quantity (I have the LCD-XC, but haven't heard the LCD-X), but very easy to listen to.
If your budget is higher and you like the Audeze sound, check out the Sony MDR-Z1R. They give my LCD-4 a run for the money in terms of bass and sound quality and are a lot more comfortable.
I forgot to mention, the Sennheiser HD800 are also super comfortable, and more refined sounding than the 600 series variants. I haven't heard the HD800s, which are supposed to be tuned a little "darker" than the HD800, but the HD800 are a different sound signature than the Audeze house sound, more neutral and detailed. Some like it, some find it a little too "analytical". I think they are great sounding headphones.