Doesn't AC (alternating current) flow in one direction for 1/2 cycle and then in the other direction for 1/2 cycle? So no matter what direction you install the fuse, isn't it going to be wrong 30 times per second?
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If it makes any difference at all, which I doubt, I would think Jea's guess is correct. However, I agree with Bob R.
Re Overhang's comment, balanced interconnects may have the shield grounded at only one end, in order to eliminate possible ground loops. That would account for directionality in the case of those cables, and the proper direction would be to have the grounded end at the source. Not sure why or if a single-ended interconnect may truly be directional (manufacturer claims notwithstanding), but in any event (considering that the shield of a single-ended interconnect serves as the signal return path, plus as an rfi/emi shield, plus as a path that may conduct current flow caused by voltage offsets between the chassis of the two components) it's a very different situation than a power line fuse.
Solid core wire is directional. The late Bob Crump
Hmm. Well, ok, I'm willing to accept their empirical findings, even though pretty much everyone in that thread admitted that the phenomenon was unexplainable.
I suspect that your bottom line about orienting from source to load is correct, in situations where it makes a difference, but I don't think that your explanation is applicable to a fuse. The fuse has no idea, so to speak, of which direction power is flowing. All it "knows" is that a current is oscillating back and forth through it, in alternate directions. It dissipates (converts into heat) an EXTREMELY small amount of power itself, equal to the square of the rms value of that current times its own very small internal resistance, but it has no way of "knowing" where the source is and where the load is.
I for one have not experimented with fuse tweaking in any of my audio equipment. My concern with audio grade fuses is how accurate is the fuse to blow under an overload condition. I have not read any actual testing of such. Of course at 30 bucks a pop who wants to.... But compared to the price of the piece of audio equipment the fuse is supposed to protect, I would consider that a small price to pay.....
I also must say I do not know, or have I checked, if the fuses have been safety tested here in the US by a credited testing lab.
There is little doubt though the fuses do make a difference in sonics. Lots of testimonials to that fact.
Direction of the Fuse? I haven't a clue.